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Genus Hoplitis Klug
[Family Megachilidae, Subfamily Megachilinae, Tribe Osmiini]
Hoplitis Klug, 1807 MFI 6: p.225; Type species: Apis adunca Panzer, 1798, monobasic.
S. No.
Name of Species with Author & Year
Distribution OR
Collection Site
1. abbreviata [Morawitz, 1875 (Osmia)]; under subgenus Kumobia Popov, 1962 Turkestan 2. acanthophora [Morawitz, 1975 (Osmia)]; under subgenus Alcidamea Cresson, 1864 Turkestan 3.
agis [Benoist, 1929 (Osmia)]; under subgenus Anthocopa Lep. & Serv.
Iran, Turkey, Griechenland
4. ahrensi [Popov, 1962 as Kumobia ahrensi]; under subgenus Kumobia Popov, 1962 Central Asia 5. alexandrina [Warncke, 1991 (Osmia)]; under subgenus Alcidamea Cresson [Zanden, 1994] Israel 6. anipuncta [Alfken, 1935 (Osmia)]; under subgenus Anthocopa Lep. & Serv. Israel, Cyprus, Kreta, Klein Asien 7. annulata [Latreille, 1811 (Osmia)]; under subgenus Annosmia Warncke Turkey 8.
anonyma [Cameron, 1908 (Megachile); Anthocopa anonyma in Michener, 1966]; under subgenus Anthocopa Lep. & Serville, 1825 Deesa 9. bicallosa [Morawitz, 1876 (Osmia)]; under subgenus Alcidamea Cresson [Zanden, 1994] Griechenland, Turkey, Caucasus 10. bifoveolata [Alfken, 1935 (Osmia)]; under subgenus Anthocopa Lep. & Serv. Israel 11. bipartita [Friese, 1899 (Osmia)]; under subgenus Anthocopa Lep. & Serv. Turkestan 12. bispinosa Zanden, 1992: p.820 as Hoplitis (Liosmia) bispinosa; under subgenus Alcidamea Cresson, 1864 Turkey 13. bytinskii [Mavromoustakis, 1948 (Osmia)]; under subgenus Anthocopa Lep. & Serv. Israel 14.
cathena [Cameron, 1908 (Megachile); Anthocopa cathena in Michener, 1966]; under subgenus Anthocopa Lep. & Serville, 1825 Bombay, Borivilli 15. caucasica [Friese, 1920 (Osmia)]; under subgenus Megalosmia Schmiedeknecht, 1885 Turkey 16. caularis [Morawitz, 1875 (Osmia)]; under subgenus Alcidamea Cresson, 1864 Turkestan 17. curvipes [Morawitz, 1871 (Osmia)]; under subgenus Prionohoplitis Tkalců, 1993 South Europe, Turkey 18. cypriaca bayburtensis Tkalcu, 1980; under subgenus Anthocopa Lep. & Serv. Turkey 19. cypriaca cypriaca [Mavromoustakis, 1938 (Osmia)]; under subgenus Anthocopa Lep. & Serv. Cyprus, Israel 20. daniana [Mavromoustakis, 1949 (Osmia)]; under subgenus Anthocopa Lep. & Serv. Israel 21. denudata [Morawitz, 1880 (Osmia)]; under subgenus Eurypariella Tkalcu, 1995 Turkestan, Mongolia upto China 22. deserti [Benoist, 1934; = tigrina Morawitz, 1972; (Osmia) in Zanden1991: p.352]; under subgenus Megahoplitis Tkalcu, 1993 Rurkey, Greece to North Africa 23. duckeana [Kohl, 1905 (Osmia)]; under subgenus Anthocopa Lep. & Serv. Turkey 24. erythrogastra [Mavromoustakis, 1954 (Bytinskia)]; under subgenus Bytinskia Mavromoustakis Turkey, Israel 25. farabensis [Popov, 1962 as Kumobia ahrensi]; under subgenus Kumobia Popov, 1962 Central Asia 26. flabellifera [Morice, 1901 (Osmia)]; under subgenus Hoplitis s.str. Syria, Israel 27. freygessneri [Friese, 1899 (Osmia)]; (under subgen. Liosmia Thomson, 1872 in Zanden, 1988); under subgenus Alcidamea Cresson, 1864 Punjab, Akalgarh: Muree (7000'), North Africa 28. fulva [Eversmann, 1852 (Osmia)]; under subgenus Megalosmia Schmiedeknecht, 1885 Turkey 29. furcula [Morawitz, 1875 (Osmia)]; under subgenus Anthocopa Lep. & Serv. Turkestan, Klein Asien 30. glasunovi [Morawitz, 1894 (Osmia)]; under subgenus Anthocopa Lep. & Serv. Turkestan 31. grossepunctata [Kohl, 1905 (Osmia)]; under subgenus Hoplitis s.str. Turkey 32. grumi [Morawitz, 1894 =jagnobensis Morawitz, 1894 (Osmia)]; under subgenus Anthocopa Lep. & Serv. Spain, Turkey, Turkestan 33. hartliebi [Friese, 1899 (Osmia)]; under subgenus Anthocopa Lep. & Serv. Israel, North Africa 34. heinrichi Zanden, 1980; (under Hoplitis s.str. in Zanden, 1992) under subgenus Chlidoplitis Griswold, 1998 Turkey 35. hemisphaerica [Alfken, 1935 (Osmia)]; under subgenus Anthocopa Lep. & Serv. Israel, Syria 36. hierichonica [Mavromoustakis, 1949 (Osmia)]; under subgenus Anthocopa Lep. & Serv. Israel 37. hofferi Tkalcu, 1977; (under subgen. Liosmia Thomson, 1872 in Zanden, 1988); under subgenus Alcidamea Cresson, 1864 Pakistan, Israel 38. holmboei (Mavromoustakis, 1948 (Osmia)]; under subgenus Hoplitis s.str. Cyprus, Rhodes 39. homalocera Zanden, 1991: p.348 as H. (Hoplitis) homalocera Jericho, & Gitit (Israel) 40. hyperplastica [Morawitz, 1894 (Osmia)]; under subgenus Alcidamea Cresson, 1864 Turkestan 41. idalia [Mavromoustakis, 1948 (Osmia)]; under subgenus Anthocopa Lep. & Serv. Cyprus, Kreta, Turkey 42. illustris Zanden, 1980; (under Hoplitis s.str. in Zanden, 1992) under subgenus Chlidoplitis Griswold, 1998 Turkey: Konya, Sertavul, Aslanko, Nevsehir, Urgup, Ankara 43.
indostana [Cameron, 1908 (Megachile); Anthocopa indostana in Michener, 1966]; under subgenus Anthocopa Lep. & Serville, 1825 Deesa 44. karakalensis [Strand, 1936: p.596 (not Popov, 1936) (Osmia)]; under subgenus Alcidamea Cresson, 1864 Iran, Turkmenistan, Pathankot 45. laboriosa [Smith, 1878; = magrettii Ducke, 1899 (Osmia)]; under subgenus Megalosmia Schmiedeknecht, 1885 Mongolia, Turkestan 46. laevifrons [Morawitz, 1872 (Osmia)]; under subgenus Alcidamea Cresson, 1864 Albania, Algeria, Quetta, Manali 47. laeviscutum [Alfken, 1935 (Osmia)]; under subgenus Alcidamea Cresson, 1864 North Africa, Turkey 48. lapidaria [Morawitz, 1877 (Osmia)]; under subgenus Hoplitis s.str. Turkey, Caucasus 49. lebanoticum [Mavromoustakis, 1955 (Osmia)]; under subgenus Alcidamea Cresson (under Micreriades in Zanden, 1988) Lebanon, Turkey 50. leiocephala [Mavromoustakis, 1954 (Osmia)]; under subgenus Anthocopa Lep. & Serv. Turkey, Israel 51. libanensis [Morice, 1901 (Osmia)]; under subgenus Hoplitis s.str. Syria, Israel 52. limassolica [Mavromoustakis, 1937 (Osmia)]; under subgenus Alcidamea Cresson, 1864 Israel, Cyprus 53. linguaria [Morawitz, 1876 (Osmia)]; under subgenus Hoplitis s.str. Turkey, Caucasus 54. lysholmi [Friese, 1899 (Osmia)]; under subgenus Hoplitis s.str. Israel, Klein Aien, Rhodes 55.
matheranensis [Michener, 1966 (Anthocopa); as Hoplitis (Anthocopa) matheranensis in Michener, 2000: p.451]; Nestings: Wain 1965 Pune, Miraz 56. meyeri [Benoist, 1934 (Osmia)]; under subgenus Alcidamea Cresson, 1864 North Africa, Israel 57. minor [Morawitz, 1877 (Osmia)]; under subgenus Alcidamea Cresson, 1864 Turkestan, Caucasus 58. mocsaryi [Friese, 1895 (Osmia)]; under subgenus Anthocopa Lep. & Serv. Southern Europe, Turkey 59. moricei [Friese, 1899 (Osmia)]; under subgenus Alcidamea Cresson, 1864 North Africa, Canary Islands to Biskara, Quetta, Alwar 60. negevensis [Warncke, 1991: LBB 23: 468-469 (Bytinskia)]; under subgenus Bytinskia Mavromoustakis Giv ot Loven, south of Paran in Wuste Negev, Israel 61. nitidula [Morawitz, 1877 = enslini Alfken, 1936, = grandior Alfken, 1935, = nursei Friese in litt., = transcaspica Morawitz, 1888 (Osmia)]; under subgenus Alcidamea Cresson, 1864 Iran, Turkestan, Caucasus 62. pallicornis [Friese, 1895 (Osmia)]; under subgenus Hoplitis s.str. Italy, Balkan Halbinsel, Turkey, Israel, Jordan 63. parasitica [Warncke, 1991: LBB 23: 484-486 (Bytinskia)]; under subgenus Bytinskia Mavromoustakis Sat-Gebirge, Hakkari, Turkei 64. peralba Zanden, 1992: p.67 as: Hoplitis (Annosmia) peralba Deesa, Syria, Egypt 65. perezi [Ferton, 1895; =papaveris convulvuli Ducke, 1899 (Osmia)]; under subgenus Anthocopa Lep. & Serv. Afghanistan, South Europe 66. persica [Warncke, 1991: LBB 23: 324-325 (Osmia)]; under subgenus Coloplitis Griswold, 1998 Chittgar, W. Teheran
[Iran]
67. pici [Friese, 1899 (Osmia)]; under subgenus Hoplitis Klug, 1807 s. str. Syria, Balkan Halbinsel 68. premordica Griswold, 1998: p. 223; under subgenus Coloplitis Griswold, 1998 Baluchistan 69. princeps [Morawitz, 1872 (Osmia)]; under subgenus Megalosmia Schmiedeknecht, 1885 East Europe, Mongolia upto China 70. rubricrus [Friese, 1899 (Osmia)]; under subgenus Anthocopa Lep. & Serville, 1825 Sahara, Biskra, Quetta, Deesa 71. ruficornis [Morawitz, 1875 (Osmia)]; under subgenus Alcidamea Cresson, 1864 Turkestan, Afghanistan 72. rufimana [Morawitz, 1875 (Osmia)]; under subgenus Hoplitis s.str. Turkestan 73. rufopicta [Morawitz, 1875 (Osmia)]; under subgenus Alcidamea Cresson, 1864 Turkestan 74. saundersi [Vachal, 1891 (Osmia)]; under subgenus Anthocopa Lepeletier & Serville, 1825 Surroundings to Mediterranean, Algeria, Quetta, Pathankot 75. sordida [Benoist, 1929 (?) (Osmia)]; under subgenus Annosmia Warncke Turkey, Israel 76. picicornis [Morawitz, 1895 (Osmia)]; under subgenus Anthocopa Lep. & Serv. Iran, Turkestan 77. recticauda [Stanek, 1869 (Osmia)]; under subgenus Hoplitis s.str. Turkey 78. semirubra [Friese, 1899 (Osmia)]; under subgenus Anthocopa Lep. & Serv. Israel, Syria 79. serrilabris serrilabris [Morawitz, 1875 (Osmia)]; under subgenus Anthocopa Lep. & Serv. Iran, Turkestan 80. simplicicornis [Morawitz, 1975 (Osmia)]; under subgenus Alcidamea Cresson, 1864 Turkestan 81. singularis [Morawitz, 1875 (Osmia)]; under subgenus Anthocopa Lep. & Serv. Turkey, Israel, Turkestan 82. stichi [Alfken, 1936 = conjuncta Alfken, 1935, not Cresson, 1864 (Osmia)]; under subgenus Hoplitis s.str. Israel 83. tenuicornis [Morawitz, 1875 (Osmia)]; under subgenus Kumobia Popov, 1962 Turkestan 84. testaceozonata [Alfken, 1935 (Osmia)]; under subgenus Hoplitis s.str. Israel 85. tigrina [Morawitz, 1871 (Osmia)]; under subgenus Megahoplitis Tkalců, 1993 Turkey, Griechenland, North Africa 86. tridentata tridentata [Dufour & Perris, 1840 (Osmia)]; under subgenus Alcidamea Cresson, 1864 Turkey, South Europe, Caucasus 87. turcestanica [Dalla Torre, 1896 = proxima Morawitz, 1875, nec Cresson, 1864 (Osmia)]; under subgenus Alcidamea Cresson, 1864 Turkestan 88. uncaticornis [Stanek, 1969 (Osmia)]; under subgenus Anthocopa Lep. & Serv. Turkey 89. unispinna [Alfken, 1935 (Osmia)]; under subgenus Anthocopa Lep. & Serv. Griechenland, Israel 90. urfensis Zanden, 1984; under subgenus Anthocopa Lep. & Serv. Turkey 91. verruciventris [Morawitz, 1886 (Osmia)]; under subgenus Alcidamea Cresson, 1864 Turkey, Caucasus 92. wadicola [Alfken, 1935 (Osmia)]; under subgenus Anthocopa Lep. & Serv. Israel 93. wahrmani batyamae Zanden, 1986; under subgenus Anthocopa Lep. & Serv. Israel 94. wahrmani wahrmani [Mavromoustakis, 1948 (Osmia)]; under subgenus Anthocopa Lep. & Serv. Israel 95. yermasoyiae asiae Tkalcu, 1979; under subgenus Anthocopa Lep. & Serv. Turkey 96. yermasoyiae yermasoyiae [Mavromoustakis, 1938 (Osmia)]; under subgenus Anthocopa Lep. & Serv. Cyprus, Griechenland
Subgeneric placements of tigrina, pici, laboriosa and curvipes amended on 11 Sept. as per suggestions of Dr. D. Yanega (California). Still to include: some species from the south-west Asia, that belong to subgenus Annosmia Warncke, Pentadentosmia Warncke, Platosmia Warncke and, Prionohoplitis Tkalců.
Political territories have been mentioned as such (as mentioned by original authors): Burma [Present day Myanmar], Ceylon [Present day Sri Lanka], Bengal [Present day East Bengal in India and the present day Bangla Desh], Punjab [western part of the state is the present day Punjab in Pakistan].
Dr. Rajiv K. Gupta
Department of Zoology
Jai Narain Vyas University
Jodhpur 342 005, India
Ph. +91-291-272 6666
E-mail: [email protected]
Visit "An Updating Bibliography of the Bees of the World" by the same author for the authors' linked references mentioned in this document.