The Origin of "anatomically modern" Homo sapiens What do we mean by "anatomically modern"? Shorter, rounder skull Vertical forehead Boney chin Smaller browridges Lighter build NOT Neandertals The earliest known specimens ~ 100-120 TYA Found in Africa Border Cave Klasies River Mouth Behavior and extent of early moderns Tool types Mostly same as Neandertals But with some innovations Blade tools Bone tools Symmetrical points More regional variation Hunting strategies More technologically advanced hunting; snares, thrown spears More varied diets; fish, shellfish, sea mammals Better knowledge of animal habits; more intense hunting and gathering Seasonal strategies Regions and sites Africa; above plus Omo, Jebel Irhud Israel; shares with Neandertals- same time or alternating? Skhul & Qafzeh ~ 100-80 TYA Europe; eventually replaces Neandertals ~30-40 TYA Vindija, Croatia Cro-Magnon, France The Great Debate The Muliregional Hypothesis Milford Wolpoff Modern peoples descended from local populations of H. erectus Gene flow maintained one, giant breeding population Adaptive changes spread, while regional quirks persisted The Population (or Complete) Replacement Hypothesis Chris Stringer Modern peoples descended from one recent population of modern H. spaiens Spread out and totally replaced archaics The Genetic (or Partial) Replacement Hypothesis Gunter Brauer Modern triats first arose in Africa Spread out & interbred with archaics Modern form more successful, archaic traits died out Evidence Fossil Earliest Moderns found in Africa, later in other regions Very few "transitional" or "hybrid" fossils In W. Europe, latest Neand. & Earliest Mod. distinct In E Europe & Isreal, possible "hybrids" Traits that Multiregionalists see continuity in, Replacementites see differences Archaeological Clear break in tool types in Europe, later Mid-east and Asia But use same tools earlier in Isreal Innovations that become pronounced later first appear in Africa Genetic Mitocrodrial DNA places ancestor all all living humans in Africa at about 100 TYA African populations have most varied DNA; means living there the longest & other pops decended from them Challanges to DNA analysis in timing and region; computer programs can give several "right" answers Modern humans are very uniform in genetic traits; not much variation between pops or individuals On the balance, a Partial Replacement seems most likely; Modern H. Sapiens arose first in Africa, then spread to other areas replacing Archaics with some amount of interbreeding.