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Cats
Cats in history
Distribution of cats
The cat as scavenger
Breeding habits
Health
Senses & Intelligence
Breeds of cats
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Cats
All cats, from large jungle cats to small house cats, belong to the CAT FAMILY, Felidae, order Carnivora.  This article discusses the domestic cat, Felis catus catus, a species indigenous to the Old World that includes such related forms as the European wildcat, F. catus sylvestris, the African wildcat, F. catus lybica, and perhaps some other distinct groups.  The domestic cat is apparently not a separate species: there is evidence of hybridization and interfertility between it and the wild races.   
The domestic form is most clearly related to the African wild race, which also ranges into areas of the Middle East. Domestic cats acquired wild-type genes wherever domestic populations were established near wild regions.  Domestic populations may therefore differ from one another because of such regional contacts.  Today, however, such genetic exchanges are limited to those few refuge areas where wild cats have not been exterminated. 
Cats in history
The domestic cat has evolved as an urban scavenger and may have originated about 8000 BC, when nomadic humans settled into village life.  From archaeological and anatomical evidence, however, it is impossible to distinguish wild from domestic types until New Kingdom times (1570-c.1085 BC) in Egypt.   
The cat was then bred by a religious cult and worshiped as a sacred animal.  This episode of cultism in Egypt has perhaps been exaggerated and, in any event, distorted in terms of its implications for the domestication process.  The cultism was undoubtedly the culmination of a long-standing familiarity between humans and cats but was not critical in bringing about the domestication of the species.  Although the cat cult was spectacular, the Egyptians domesticated many species, from snakes to gazelles, and the cat may simply have been bred to protect granaries from rodents. 
The spread of domestic cats appears to have followed the pattern of progressive urbanization.  The appearance of local domestic cat populations therefore correlates with the spread of civilization from the Middle East, and domestic cats now inhabit every continent except Antarctica and most of the world's oceanic islands.  Cats have reverted to a wild state in many habitats where they were not originally found, and in some places they persist even though humans have abandoned the area. 
Distribution of cats
Recent studies indicate that cat populations range from sparse to more than 2,000 per square mile, depending on several factors.  The largest populations exist in urban areas, especially in so-called Mediterranean climates, where opportunities abound for year-round food and shelter.  Only in certain regions such as Western Europe and North America are cats kept as pets in numbers sufficient to support such large, subsidiary industries as veterinary services, pet foods and novelty products, welfare societies, and publications. Elsewhere, cats are tolerated but not regarded as pets.  Their impact on urban ecology is difficult to judge, and basic studies to assess their role in vermin control and as disease vectors are needed. 
The cat as scavenger
Apart from a decrease in body size, the most notable anatomical contrast between domesticated and wild cats is the increased intestinal length in the former.  This may correlate with the cat's turning from chiefly predatory to chiefly scavenging habits, and the resulting reliance on a low-protein diet. Theoretically, increased small-intestinal length would maximize absorption of amino acids from a minimum protein diet.   
Domestic cats, like their ancestors and contemporary relatives, are twilight hunter-scavengers.  The urban domestic cat has largely shifted toward scavenging behavior.  One study of the stomach contents of stray city cats revealed that only 3 percent of their diet comprised rodents;  the balance was garbage.   
Many studies have been devoted to the question of the depredations of cats on birds.  Cats would seem to have only a modest effect, and generalizations are risky. 
Breeding habits
Domestic cats, like their wild relatives, are seasonally polyestrous;  they breed more than once a year, depending on conditions.  The breeding cycle is controlled by sensitivity to light.  At equatorial latitudes, where there are longer periods of daylight, cats may breed throughout the year and have three or four litters.  In temperate latitudes, they normally have two litters a year, one in the early spring and another in late summer.   
Domestic cats living under artificial light may breed at any time.  Reproductive life begins at 7.5 months and continues until an advanced age--15 years or more.  The gestation period is usually 63 days, and the average litter is four kittens, except for the usually smaller first litter, and litters born to older females (8 years of age or over).
Health
A cat's normal body temperature is 38.6 deg C (101.5 deg F), with variations of 0.5 deg C (1 deg F) above or below.  The critical environmental temperature for the cat is about 36 deg C (97 deg F), at which point its body temperature rises and the cat begins to pant to cool itself.  Cats have sweat glands both in the pads of their feet (eccrine glands) and scattered over their bodies (apocrine glands), but sweating is not a temperature-regulating mechanism in cats. 
One of the most common and destructive cat diseases is feline viral rhinotracheitis, commonly called FVR or cat flu.  It is not contagious to humans or other animals.  An infected cat develops a fever in two to five days, begins sneezing and drooling, and becomes lethargic.  This is followed by a heavy discharge from the eyes and nose, which tends to clog the nostrils and paste the eyelids shut.  Treatment consists of antibiotics to counter secondary bacterial infections, cleansing the eyes and nose, and administering fluids to offset dehydration.  Feline distemper, also called panleukopenia, feline enteritis or gastroenteritis, or cat fever or plague, is a common, highly contagious viral disease characterized by its explosive course: death may occur as soon as eight hours after the first signs of illness appear. The virus is transmitted by even brief contact with contaminated material. Dogs are not affected.   
Fleas are tiny, dark red, blood-sucking insects that may be seen moving quickly through the cat's hair.  Their droppings resemble particles of black sand, and the presence of this gritty substance indicates an infestation of fleas, even if the fleas themselves are not seen.   
Fleas can be removed by special shampoos, by powdering with rotenone insecticide, or by the wearing of flea collars (if the cat is not allergic).   
Roundworms are slender, whitish worms, 2.5 to 10 cm long (1 to 4 in), with tapered ends. Heavy infestation, especially in kittens, causes coughing, vomiting, potbelly, bad breath, and stunted growth.  Roundworms can be transmitted to humans. Treatment consists of administering piperazine tablets.   
As with other pets, cats that roam outside the house can bring in parasites and transfer them and the diseases they may carry to humans.  Although studies of cats as disease vectors are needed, few of the diseases appear serious;  a few cases of BRUCELLOSIS and, extremely rarely, of plague have been reported.  Pregnant women are cautioned about handling cats and their litter boxes because of the possibility of transmitting TOXOPLASMOSIS to their fetuses;  the eventual effects of the disease when contracted before birth can be quite serious. Cats may also transmit such infestations as ringworm and roundworm, and some people exhibit a severe allergic reaction to the animals--most likely, it has been suggested, to dried spittle on their fur.  Cat-scratch disease, a benign but sometimes painful disease of short duration, has recently been found to be caused by a bacillus;  despite its name, the disease may be transmitted by many kinds of scratches besides those of cats. 
Senses & Intelligence
Like all felids, domestic cats have senses evolved to facilitate twilight predation.  Thus, they are especially well equipped to detect slight movements in semidarkness.  The field of visual overlap approximates that found in primates and gives cats full depth perception.Cats are also extremely sensitive to sounds, including frequencies beyond the range of human hearing.  Sense of smell is not greatly elaborated in felids, although there is some evidence that olfactory cues are used in courtship behavior.   
Domestic cats are apparently no more or less intelligent than wild cats. They may be more versatile than their wild relatives because of adaptations to urban life, but, originating from basically solitary ancestors, they are not as tractable or malleable as other domesticated species.  Many of the apparent differences between cat and dog behavior, for example, stem from motivation rather than innate ability. Under controlled behavioral experiments, cats readily perform highly complex tasks.
Breeds of cats
The cat differs strikingly from other domesticated animals, in that it has not, until recently, been bred selectively.  Hence, most of the observed genetic changes and variations in cats probably result from inherent adaptations to domesticity or to the perpetuation of novel anomalies.  Most of the better-understood genetic variations in cats relate to differences in color, pattern, and texture of the coat, and to such minor skeletal peculiarities as taillessness or supernumerary digits.  Cats are remarkably similar in shape and size.   
The so-called breeds of cats are normal variants in free-ranging populations that do not transmit their traits generation after generation like pure strains.  The Manx cat did indeed originate from the Isle of Man as the result of a local 17th-century mutation, but it does not breed true.  There is no evidence that ABYSSINIAN cats are of Ethiopian origin, and PERSIAN cats are actually more common in the Soviet Union than in Iran.  The origin of most of the older so-called breeds appears to be late 19th-century and early 20th-century England, despite the fanciful histories that are frequently proffered. 

Copyright © 1999 Sasha Grinshpoon.
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